1.Dark-field, multiple-exposure photographs of live spermatozoa of a number of echinoderms were analysed. 2. Bends develop at the base in pairs, as they do in flagella with symmetrical waveforms. However, the angles of these bends do not cancel, so that microtubular sliding-of up to over 50% of that associated with bend propagation-is transferred distally. This sliding implies that (a) microtub...