Few microorganisms match the impact that the oomycetes have had on mankind. This distinct lineage of eukaryotes is wellknown for its most notorious member, Phytophthora infestans, the agent of the nineteenth century Irish potato famine, and several other devastating pathogens of cultivated and wild plants [1]. Indeed, more than 60% of oomycete species infect plants [2]. Less known is the fact t...