We agree with Sandve et al. (2015) that the nomenclature of Aegilops /Triticum lineages is complex. Indeed, there is a contradiction in how they themselves have defined ‘theDgenome lineage’ in their Letter (Sandve et al., 2015), compared to their earlier paper (Marcussen et al., 2014). Fig. 1 of Sandve et al. (2015) defines it as a clade comprising D + S* + M genome species; this definition is ...