R.A. Adams and S.C. Pedersen (eds.), Bat Evolution, Ecology, and Conservation, DOI 10.1007/978-1-4614-7397-8_12, © Springer Science+Business Media New York 2013 Abstract To be evolutionarily stable, cooperative behavior must increase the actor’s lifetime direct fi tness (mutualism) or indirect fi tness (altruism), even in the presence of exploitative, noncooperative “cheaters.” Cooperators can ...