A hymenial cell initial was binucleate in Coprinus cinereus but uninucleate in Volvariella bombycina. Paraphyses and facial cystidia remained binucleate in C. cinereus, but marginal cystidia were the apical cells of branches from the multinucleate gill trama. becoming swollen when gills pulled away from their connection to the stipe, and were multinucleate. In V. bombycina. all mature cystidia ...