Rhynchocephalia, a once diverse clade of lizard-like reptiles, had nearly global distribution during much the Mesozoic. By Late Cretaceous rhynchocephalians underwent marked reduction in their diversity and biogeographic range, with sparse records only deeply nested taxa (e.g., Sphenodontinae) found Upper Cretaceous–Paleocene Patagonia Miocene–Recent New Zealand. Here we describe partial dentar...