throcyte antigen loci covering the whole genome. De-regressed breeding values were available for 29 traits, including 20 conformation, 2 fertility, 4 birth, 2 workability, and 1 longevity trait. After determining the most likely marker haplotypes for all grandsires based on the genotypes of their sons, multi-marker regression analysis of the trait data was used to scan the genome for QTL. Chrom...