ALTHOUGH IT HAS BEEN FORMALLY RECOGNIZED since the end of the 19th century that invertebrates can learn (Romanes 1895), the modern neurobiological analysis of invertebrate learning did not begin until the 1960s. Starting in that decade, pioneering investigators began to use intracellular electrophysiology to probe the basic mechanisms of learning in higher invertebrates (Bruner and Tauc 1966; K...