Many economists have observed that wealth is systematically lower in the tropics than elsewhere (Sala-i Martin, 1997; Nordhaus, 2006). Determining why this is remains a major puzzle. Leading hypotheses include, inter alia, the tropics’s disease burden (Gallup, Sachs and Mellinger, 1999), biota available for domestication (Diamond, 1997), distance from trading partners (Frankel and Romer, 1999),...