Eriophyoid mites (Acariformes, Eriophyoidea) are microscopic chelicerates morphologically greatly preadapted to endoparasitism on plants. Members of at least six phylogenetically distant eriophyoid genera from two families homoplastically transitioned and acquired the ability penetrate under plant epidermis feed parenchymatous cells, usually causing necrosis. Theoretically, endoparasites expect...