a. alizadeh department of plant protection, college of agriculture and natural resources, university of tehran, karaj m. javan-nikkhah department of plant protection, college of agriculture and natural resources, university of tehran, karaj, iran r. zare department of botany, iranian research institute of plant protection, agricultural research, education and extension organization (areeo), tehran, iran kh. b. fotouhifar department of plant protection, college of agriculture and natural resources, university of tehran, karaj, iran u. damm senckenberg museum of natural history görlitz, germany e. h. stukenbrock environmental genomics, botanical institute, christian-albrechts university of kiel