maryam yazdanian hepatitis and aids department, pasteur institute of iran, tehran, ir iran. tel/fax: +98-2166969291 arash memarnejadian hepatitis and aids department, pasteur institute of iran, tehran, ir iran. tel/fax: +98-2166969291 ; hepatitis and aids department, pasteur institute of iran, tehran, ir iran. tel/fax: +98-2166969291 mehdi mahdavi virology department, pasteur institute of iran, tehran, ir iran. tel/fax: +98-2166496682 seyed mehdi sadat hepatitis and aids department, pasteur institute of iran, tehran, ir iran. tel/fax: +98-2166969291 fatemeh motevali hepatitis and aids department, pasteur institute of iran, tehran, ir iran. tel/fax: +98-2166969291 rouhollah vahabpour hepatitis and aids department, pasteur institute of iran, tehran, ir iran. tel/fax: +98-2166969291

Viraj Kulkarni Margherita Rosati Rashmi Jalah Brunda Ganneru Candido Alicea Lei Yu Yongjun Guan Celia LaBranche David C Montefiori Alan D King Antonio Valentin George N Pavlakis Barbara K Felber

seyed mansour razavi department of community medicine, tehran university of medical sciences, tehran jamshid hadjati hadjati department of immunology, tehran university of medical sciences, tehran payman salamati sina trauma and surgery research center, tehran university of medical sciences, tehran

seyyed meysam abtahi froushani assistant professor of immunology, department of microbiology veterinary medicine faculty, urmia university, urmia, iran hadi esmaili gouvarchin galee msc student of immunology, department of microbiology, veterinary faculty, urmia university, urmia, iran mahsa khamisabadi student of veterinary medicine, veterinary faculty, urmia university, urmia, iran. bita lotfallahzade student of veterinary medicine, veterinary faculty, urmia university, urmia, iran.