somayeh ghasemian department of biology, college of sciences, shiraz university, shiraz, iran. hamid reza esmaeili department of biology, college of sciences, shiraz university, shiraz, iran. mohsen nokhbatolfoghahai department of biology, college of sciences, shiraz university, shiraz, iran. abdolrahim pazira department of fisheries, islamic azad university, bushehr branch, bushehr, iran.